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Research Lab Bench
Karol Subczynski

W. Karol Subczynski, PhD, DSc



  • Biophysics
    MFRC 2008

Contact Information


DSc, Natural Sciences with specialization in Biophysics, Jagiellonian University, Krakow, Poland, 1984
PhD, Physico-Mathematical Sciences with specialization in Biophysics, Lomonosov Moscow State University, Moscow, USSR, 1976
MSc, Physics, Lomonosov Moscow State University, Moscow, USSR, 1969


Shortly after completing my training, I joined the Biophysics Department, Institute of Molecular Biology, at Jagiellonian University, Krakow, Poland, as a teaching/research assistant. I joined the faculty in 1977; in 1995, the President of Poland awarded me the title of Professor of Biophysics. From 1988–1991, I was Chair of the Biophysics Department, and from 1992–2000, I was Head of the Laboratory of Structure and Dynamics of Biological Membranes. I first visited Milwaukee in 1980; since then, I have been at the National Biomedical EPR Center about 50% of the time. In 2000, I emigrated to the United States and joined the MCW faculty as an Assistant Professor of Biophysics; in 2002, I became an Associate Professor of Biophysics; and, in 2010, I became a Professor of Biophysics.

I have directed 14 graduate students pursuing their MSc degrees and supervised five students pursuing their PhD degrees.

I have coauthored more than 150 original papers, reviews, and book chapters.

Research Interests

北方今冬再战雾霾:多行业错峰生产 山东水泥企业停产

  • 'Fruitvale Station' represents another remarkable feature debut, by the director Ryan Coogler. Its subject matter-the fatal shooting of Oscar Grant, a black, unarmed 22-year-old by a white transit cop in Oakland, Calif.-might have made a documentary. Mr. Coogler had the better idea of exploring it as a dramatized, and to a considerable extent fictionalized, account of the day in Oscar's life that preceded his death. He's played by Michael B. Jordan, whose performance can break your heart or fill it with joy, sometimes simultaneously.
    单词expansion 联想记忆:
    The list has four British universities, which are Cambridge, Oxford and London's University College and Imperial College. The University of Tokyo is also included in top 20.
  • 开发商:Frontier、Microsoft Studios
    A lone European telephone bidder paid a top price of 9.1 million for a smaller 1825 version of John Constable’s admired but difficult-to-love 1824 vertical landscape, “The Lock,” that had been in the same British family since 1855.
    8. Julia Roberts - $12 million
    在电影《007:大破天幕杀机》中饰演邦女郎的贝纳尼丝·玛尔洛(Berenice Marlohe),已经确认加盟导演特伦斯·马立克(TerrenceMalick)执导的最新电影作品,目前此影片正在奥斯汀进行拍摄。
  • ['t?nl]
    The awards, announced during a ceremony in Amsterdam on March 14, are based on millions of international passenger surveys.
  • 节目5 创意器乐演奏《琴筝和鸣》雅尼(美国)、常静等
    1. “The Assassin” and “Mad Max: Fury Road” (tie) Hou Hsiao-Hsien and George Miller directed the year’s two best commercial movies, both of which should be seen on the biggest screen you can find.
    你是否一直有一个作家梦,但又不知道从何着手?那么你是否考虑过自助出版呢?感谢亚马逊,这种想法已经成为可能。亚马逊的Kindle Direct Publishing工具可以让你自行制作电子书,CreateSpace能够帮助制作纸质书籍,而ACX则是有声电子书的制作部门。我在出书时曾使用过这三款软件。
    Will the S&P 500 finish the year above 2,650
    8. Illiteracy
  • 成功的CEO都保持着都极有规律的生活,他们每天早起,锻炼身体,身边会放一些点心补充能量,每周拿出几个晚上进行社交。
  • 11月,中国的出口总值增速是预期的2倍多,这增加了中国的贸易顺差。
    No. This crisis hit Amsterdam in 1772, after a respected Dutch investment syndicate made a disastrous bet on shares of the British East India Company.
    China is the most striking wild card that will drive agricultural markets, said Stefan Vogel, head of agri commodity markets at Rabobank,

Research Projects

  • Lipid Rafts in Eye Lens: Discrimination by Pulse EPR is an R01 award by the National Eye Institute (December 1, 2004–November 30, 2009).
  • Cholesterol Crystalline Domain Function in Eye Lens: EPR Spin-Labeling Studies is an R01 award by the National Eye Institute (December 1, 2009–May 31, 2015).
  • Is Cholesterol Crystalline Domain a Barrier to Oxygen Transport? is a grant award by the NIH Fogarty International Center (February 1, 2009–December 31, 2012).
  • Lipid Domains in Lens Membranes of a Single Eye: EPR Spin-Labeling Studies is an R01 award by the National Eye Institute (June 1, 2015–May 31, 2020).

Lab Members
Natalia Stein, PhD, Postdoctoral Fellow

Dr. Akihiro Kusumi, Institute for Frontier Medical Science, Kyoto University, Kyoto, Japan
Dr. Marta Pasenkiewicz-Gierula, Biophysics Department, Institute of Molecular Biology, Jagiellonian University, Krakow, Poland
Dr. Marija Raguz, Department of Medical Physics and Biophysics, University of Split School of Medicine, Split, Croatia
Dr. Justyna Widomska, Department of Biophysics, Medical University of Lublin, Lublin, Poland
Dr. Alexander N. Tikhonov, Department of Biophysics, Faculty of Physics, Lomonosov Moscow State University, Moscow, Russia

Recent Book Chapters

Subczynski W.K., Widomska J., Mainali L.妮诗原来就读于奥顿肖杨树街小学,现在就读于威辛顿女子学校,今年九月会进入奥尔特林厄姆女子语法学校。她与父母尼朗加和施若咪一起生活, 2001年,一家三口从斯里兰卡搬到了曼彻斯特。44岁的父亲尼朗加称赞妮诗在杨树街小学的启蒙老师们功不可没。他说:“我们很早就知道她有天赋,她很小就开始阅读和写字,而且非常擅长数学计算。我们让她做一些有挑战又有趣味的事。作为父母,你不想让孩子的天赋荒废掉,但是在他们的童年时期又需要平衡。她只是一个普通的10岁小孩,真的就像任何其他的10岁小孩。她喜欢阅读、骑车、散步,我们都为她感到骄傲。”
, in: Oxygen Transport to Tissue XXXIX, Halpern H., LaManna J., Harrison D., Epel B., Eds. Advances in Experimental Medicine and Biology, vol 977. (Springer, Cham), 2017, pp. 27-34.

Subczynski WK, Wisniewska A, Widomska J. Xanthophyll–Membrane Interactions: Implications for Age-Related Macular Degeneration, in: Carotenoids and Retinal Disease, Landrum JT and Nolan J, Eds. (CRC Press), 2014, pp. 203-221.


  • La La Land producer Jordan Horowitz returned to the microphone after being told about the mix up and said 'Moonlight won Best Picture' and insisting that 'this is not a joke'.

    (Subczynski WK, Pasenkiewicz-Gierula M.) Cell Biochem Biophys. 2020 Sep;78(3):241-247 PMID: 32602057 PMCID: PMC7403164 07/01/2020

  • This journey into the mind and feelings of an 11-year-old-girl may be Pixar’s wildest adventure yet. It’s a very funny workplace sitcom (with exuberant, touching performances from Amy Poehler, Phyllis Smith, Mindy Kaling and others), an ingenious allegory of psychological development, and an almost unbearably moving and honest defense of the role of sadness in our lives. (Read the review.)

    (Boban Z, Puljas A, Kovač D, Subczynski WK, Raguz M.) Cell Biochem Biophys. 2020 Jun;78(2):157-164 PMID: 32319021 PMCID: PMC7608754 04/23/2020

  • 13 Reasons Why

    (Widomska J, SanGiovanni JP, Subczynski WK.) Nutrients. 2020 May 07;12(5) PMID: 32392888 PMCID: PMC7284714 05/13/2020

  • ['pl?tf?:m]
    An investigation is continuing into how La La Land was wrongly named Best Picture rather than the triumphant Moonlight.

    (Mainali L, Pasenkiewicz-Gierula M, Subczynski WK.) Curr Eye Res. 2020 02;45(2):162-172 PMID: 31462080 PMCID: PMC6980519 08/30/2019

  • Video editors likely benefited from the same factors that caused the increase in photography jobs. It stands to reason that this kind of work expands alongside the need for increasingly sophisticated and appealing website designs.

    (Raguz M, Kumar SN, Zareba M, Ilic N, Mainali L, Subczynski WK.) Cell Biochem Biophys. 2019 Dec;77(4):309-317 PMID: 31625023 PMCID: PMC6858498 10/19/2019

  • (实际上,在过去三年的节日假期里,捷蓝航空公司绝对不是航班起飞延误最糟糕的航空公司。这项殊荣要授予Comair公司,这是达美航空公司经营地区航线的公司。不过该公司在2012年假日季节开始前停业,因此我们没有让这家公司上榜。)
    单词response 联想记忆:

    (Stein N, Mainali L, Hyde JS, Subczynski WK.) Appl Magn Reson. 2019 Jul;50(7):903-918 PMID: 31244509 PMCID: PMC6594395 06/28/2019

  • Put succinctly, there was no way to describe or explain the crosscurrents of 2014. And I’m not convinced that the benefit of hindsight will make what happened this year any clearer years from now.

    (Widomska J, Subczynski WK.) Nutrients. 2019 May 15;11(5) PMID: 31096723 PMCID: PMC6566707 05/18/2019

  • 在赛场上,绰号Baby的天才逃亡车手(安塞尔?埃尔格特饰)依靠专属的音乐节奏开车而独占鳌头。与梦中女神(莉莉?詹姆斯饰)邂逅之后,Baby看到了希望,想与以往不堪的生活断个干干净净。Baby被迫为黑社会老大(凯文?史派西饰)干活,在派给他的最后一单中,危机一触即发。面前是巨大的挑战,对他的生活、爱人和自由来说,形势都岌岌可危……

    (Tikhonov AN, Subczynski WK.) Cell Biochem Biophys. 2019 Mar;77(1):47-59 PMID: 30460441 PMCID: PMC6524781 11/22/2018

  • 公司:网络投票公司Everyone Counts
    单词commercial 联想记忆:

    (Mainali L, O'Brien WJ, Subczynski WK.) Exp Eye Res. 2019 01;178:72-81 PMID: 30278157 PMCID: PMC6361697 10/03/2018

  • 该车票系统由中国铁道科学研究院电子计算技术研究所研发。该研究所副所长朱建生称:“如果所有这些车票排列起来,可以绕地球7圈。”

    (Widomska J, Subczynski WK.) Exp Eye Res. 2019 01;178:238-246 PMID: 29908882 PMCID: PMC6447089 06/18/2018

  • 6. 2012年搞笑诺贝尔奖之文学奖

    (Plesnar E, Szczelina R, Subczynski WK, Pasenkiewicz-Gierula M.) Biochim Biophys Acta Biomembr. 2018 Feb;1860(2):434-441 PMID: 29079282 PMCID: PMC5732854 10/29/2017

  • It’s ironic that in a country as populous as China, a meme about loneliness would be so popular. This seemingly irrational line has been repeated in every possible permutation, on every possible occasion, e.g. “What big brother is reading not a book, but loneliness.”

    (Mainali L, Camenisch TG, Hyde JS, Subczynski WK.) Appl Magn Reson. 2017 Dec;48(11-12):1355-1373 PMID: 29805201 PMCID: PMC5967259 05/29/2018